Mixed conifer and beech forests complement each other in the use of resources, unless rainfall is low

Complementarity between Scots pine and broad-leafed species in the use of available resources such as water may increase the growth of mixed forests comprising both species compared with pure forests with only one. However, the lack of rainwater reduces this advantage in species that, like the Scots pine, do not tolerate shade, because increased competition for water does not allow them to compete for light.