A team of wildlife researchers from the U.S., South Africa and Eswatini (formally known as Swaziland) has found via experiments that ungulates (hooved animals) living in game reserves in Eswatini fear elephants as much as they do leopards. In their study, reported in the journal Biology Letters, the group played animal noises over speakers and recorded the reactions of the ungulates.
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Source: Phys.org